• pradudruvuḥ — they fled
  Plural perfect of pra + √dru (to run). The verb of flight. The Bhāgavatam uses it for Kṛṣṇa's enemies again and again — when the front rank falls, the rest run. The doctrine: demoniac collectives have no internal cohesion.
• prāṇa - parīpsavaḥ — desirous of saving their lives
  Compound: prāṇa (life-breath) + parīpsu (desiderative of √āp, 'wishing to obtain'). Lives become the only object once power has been broken. Gauḍīyas note: the demon, denied dominance, instantly becomes a mere body wanting to keep its breath.
• śeṣāḥ — the remaining ones
  Plural of śeṣa — what is left over. The remnant. The five chiefs being named makes the unnamed simply 'remainders' — the Bhāgavatam will not catalogue cowards individually.
• mallāḥ — the wrestlers
  Plural of malla. The class of professional wrestlers — paid champions of the king. The poet groups them by class as soon as they lose their leaders; demoniac identity disappears into anonymity once the head is removed.

The five named wrestlers dead, the rest run for their lives — like soldiers who, the moment the standard-bearers fall, remember they are mortal. Demoniac courage was always a borrowed courage; when the borrowers fall, the lenders flee.